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It all started with two things happening, God disappearing and humans creating a new type of war machine.
These new machines had the ability to use blood as fuel, making them very efficient on the battlefield. In the game you play as V1, a special type of war machine that has an exoskeleton that allows it to siphon blood directly through it's armor, making it very dangerous since it can recover from damage very fast. But shortly after the creation of V1 a new peace was founded and all war machine production was halted, meaning only one V1 model was ever made, A V2 model was (for unknown reasons) made after the new peace which couldn't siphon blood but was made to be tougher due to the lack of blood.
Humanity also discovered Hell (which might have directly led to the new peace) giving war machines a new purpose, exploring hell and its layers.
But there was now a shortage on blood since no bloodshed was needed, so there was eventually a robot uprising leading to the extinction of the human race. Since most humans were sinner in one way or another the majority of them went to hell, which is why the machines made their way down there as well.
While all of this is happening God has suddenly disappeared, causing a couple thing to happen, Heaven is in chaos and is trying to piece itself back together, down in the lust layer king Minos is making a reform and allowing the citizens of the lust layer to live a decently normal life in the now massive city Minos created.
In the greed layer king Sisyphus is making an insurrection against heaven, but after the holy council was established he was quickly shut down. The supreme angel Gabriel was assigned the task of killing Minos and returning the lust layer to its former "glory". Minos made no attempt to fight Gabriel trying to reason with him and was struck down, his soul was then imprisoned inside his now infested corpse and Gabriel took on the role of judge in Hell.
The game starts with you trying to find the gates of Hell, on the way there you find a gun and while moving further down you start seeing another machine that's killing demons in a similar way to how you were up until this point. After a couple more sightings you finally meet face to face, you end up killing the machine and taking its shotgun. Finally you arrive at the gates of Hell and on the door there is a text: "abandon hope all ye who enter here", the statues beside the door come alive and you have to face them, they are called Cerberus despite their human shape because they act as protectors throughout Hell.
After defeating them and going through Hells gates you arrive in Limbo, a purgatory like plain where everything seems peaceful but once you look closer it becomes clear that this place is nothing but an illusion, the sky is nothing but a screen and the bird noises and calm music is coming from speakers behind trees. You make your way through limbo and find a big door that needs three skulls to be opened, while finding them you see a red version of yourself zooming past a doorway. Once the door has been opened and you walk in V2 flies through a window and lands gracefully, it greets you before attacking. This is one of the toughest bosses for new players and is comparable to Bladewolf from Metal Gear Rising in that it forces you to play the game in the intended way and acts as a final exam for if you understand all of the games mechanics. After defeating and almost killing V2 it flees to repair itself leaving its right arm behind, you pick it up and learn that it can work as a shockwave to push back enemies.
You enter the lust layer and see how it has been brought to ruin by the angels, in the distance you can see the corpse of king Minos looking over the trampled city with spotlight like eyes looking for sinners to punish. With every level you move towards the corpse, getting closer and closer until he's right above you, you use the old subway to move forward but you are suddenly stopped by the corpse and you have to fight him, when you get him down to half health two parasites pop out of his eyes and start shooting at you, showing that he is nothing more than a puppet.
After killing him you descend down into the gluttony layer, a place made of meat, bones and eyes, you hear a voice telling you to "turn back or else you will be crossing the will of god", he tells you that he is the righteous hand of the father and that he will "rend you apart", of course you ignore this and keep moving until you reach an arena with a beating heart behind it. From a beam of light Gabriel, the judge of Hell appears and tries to kill you for trespassing, after you kill him he goes into shock wondering how a mere machine could best a supreme angel, he teleports away and assures you that this isn't over. As punishment for loosing against a machine Gabriel is given 24 hours before the light is ripped from him, killing him.
You keep moving, entering the golden desert of greed where the Sisyphean Insurrectionists are forced to haul boulders up famous landmarks for eternity as punishment for their attempt to overthrow heaven. you end up killing one of them and descending into a pyramid where you then climb to the top to fight V2 for the second time, its got more weapons and a new arm this time, after defeating it again it tries to run away just like last time but you case after it, sliding down the pyramid you kill V2 for good. stealing its arm once more. this time it's a grappling hook that allows you to pull small enemies towards you and you towards big enemies.
In the wrath layer, the raging sea of Styx you make your way towards a big ferry where you fight and kill the Ferryman to gain entrance to the ferry, finding a rocket launcher inside and seeing a giant sea serpent that later capsizes the ferry forcing you to find a way out. You end up on the ocean floor where you see the silhouette of the sea serpent, you make your way up to the surface where you fight the serpent to descend into the heresy layer.
In heresy you meet Gabriel again, this time his pride isn't in the way of him using his swords splendor and justice, and he's pissed at the fact that he lost so when you defeat him again he starts to question the council and its power. After many hours of thinking he decides to kill every single one of the council members, even if that means he won't be able to regain his holy light. He displays the severed head of a council member to the crowd outside to send the message that he was now in control, now enlightened he will face V1 once more in chapter 3 (which has yet to release).
But there's more, in gluttony there is also a massive door that requires you to get perfect rank on all levels to enter, behind this door is a giant flesh prison which once killed, opens up to reveal Minos prime. A prime soul is an incredibly rare occurrence in which a soul amasses so much power that it no longer requires a Husk as a vessel to manifest physically. As manifestations of pure will, prime souls are incredibly powerful, to the point that even the prideful angels see them as a threat and will use any means necessary to stop them from forming. Minos prime thanks you for freeing him but also tells you that the crimes you have committed against humanity deserve the death sentence, Minos Prime is an incredibly powerful enemy, capable of chaining attacks to kill V1 in seconds. It only takes a moment of inattention to be utterly destroyed. After many failed attempts you finally beat him, and your reward is a log by the humans from when they were still alive. The log talks about the terminals and elevator shafts in the game used to transition between levels, it explains how they work almost as teleporters, disassembling whatever is put in at the molecular level to then transport it somewhere else, it talks about how the first successful attempt of using these transporters was a vinyl of the song "were you foolin" by Richard Himber, though it never seems to play the vocal part of the song, some claim to have heard it but had no recording equipment on them, so it is still not confirmed.